Tracker is the story of Daggon, an alien life form who lands on Earth from the planet Cirron on a mission to recapture 218 prisoners who had escaped from the planet SAR TOP in the Migar Solar System in the form of "life forces," which then took over various human identities. He has a device to capture the life forces and contain them in spheres, after which he is to return them to SAR TOP. He lands in an abandoned field in the outskirts of Chicago where he takes on the form of an underwear model and adopts the name "Cole" from an underwear ad he sees on a billboard. He later meets Mel Porter, a Chicago bar owner with an outgoing British bartender that she inherited along with her grandmother's police bar. Though initially wary, Mel gradually comes to accept Cole and lets him stay, offering him some of her ex-boyfriend's clothes.
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